| 1972-1976 | A.B., Chemistry | Dartmouth College |
| 1976-1983 | M.D./Ph.D., Human Genetics | University of Michigan |
| 1983-1988 | Postdoctoral Fellow, Molecular Neurobiology | California Institute of Tehcnology |
| 1988-1994 | Assistant Professor, Microbiology & Molecular Genetics | University of California, Irvine |
| 1994-1998 | Associate Professor, Microbiology & Molecular Genetics | University of California, Irvine |
| 1998-Present | Professor, Microbiology & Molecular Genetics | University of California, Irvine |
| 2004-Present | Professor, Anatomy & Neurobiology | University of California, Irvine |
| 2008-Present | Associate Dean for Graduate Students, School of Medicine | University of California, Irvine |