| 1985 | Ph.D. | University of California, San Diego |
| 1986-1989 | Postdoctoral Work | Stanford University |
| 1989-1994 | Assistant Professor, Development & Cell Biology, Anatomy & Neurobiology | University of California, Irvine |
| 1994-2001 | Associate Professor, Development & Cell Biology, Anatomy & Neurobiology | University of California, Irvine |
| 2001-Present | Professor, Development & Cell Biology, Anatomy & Neurobiology | University of California, Irvine |
| 2006-2014 | Professor, HHMI-UCI Professor Program | University of California, Irvine |