| 1973 | B.S., Biological Science | University of California, Irvine |
| 1978 | Ph.D., Biological Sciences | University of California, Irvine |
| 1978-1979 | Postdoctoral Work, Psychobiology | University of California, Irvine |
| 1980-1981 | Postdoctoral Work, Dept of Neurobiology | SUNY at Stoneybrook |
| 1981-1986 | Assistant Professor, Dept of Anatomy | University of California, Irvine |
| 1986-1992 | Associate Professor, Dept of Anatomy | University of California, Irvine |
| 1992-Present | Professor, Dept of Anatomy & Neurobiology, Dept of Neurobiology & Behavior | University of California, Irvine |
| 2015-2017 | Chair (Interim), Dept of Anatomy & Neurobiology | University of California, Irvine |
| 2017-Present | Chair, Dept of Anatomy & Neurobiology | University of California, Irvine |